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March 8
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Megalosaur interrupted by ScottHartman Megalosaur interrupted by ScottHartman
As a general rule most books sort of gloss over the transition from basal coelophysoid or ceratosaurian grade theropods to allosaurids, but megalosaurs are actually a fascinating bunch. They not only rose to prominence during the Middle Jurassic, but they continued to compete in the Northern Hemisphere alongside sophisticated allosauroids, and managed an impressive Cretaceous radiation (as spinosaurids). Sadly a lot of these specimens are not terribly complete, so if we want to restore them we need to do a lot of gap filling.

Torvosaurus is actually surprisingly complete once all of the referred specimens are accounted for (mercifully they also have quite a bit of overlap between material, so cross-scaling isn't such a challenge). So perhaps not surprisingly that's what was used to help fill out Megalosaurus and the new T. gurneyi.

Marshosaurus is too far from Torvosaurus to use it, so instead Condorraptor and Piatnitzkysaurus stood in. Also, to be fair to the megalosaurs and the completeness issue some of the more basal taxa like Eustreptospondylus and Piatnitzkysaurus are more complete, but I haven't got to them (yet!).
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Ladymedusa218 Featured By Owner Jul 28, 2014  Hobbyist General Artist
megalosaurs descended into spinosaurs??? wow ive never heard of that!!!what "links" would be between these two?? also is it thought that maybe the bones of megalosaurus may actually be juvenile torvos??? well i guess actually it would be megalosaurus that would be the correct genus name since it was named first. id love to know the details about all of this :) 
ScottHartman Featured By Owner Jul 29, 2014  Professional Digital Artist
Since I'm sort of swamped atm I'll point you to the appropriate Wikipedia page:… Not only does it have some fairly recent family trees, but if you meander down towards the works cited at least one paper is available to download (and it has all the characters).

The easiest not-so-technical characters to see are a very large thumb claw, and an elongated head (sure, it's taken further in spinosaurids, but megalosaurs also have long heads).
Asuma17 Featured By Owner Jul 17, 2014  Hobbyist General Artist
Come to pass the Torvosaurus seems to pretty well complete though I still can't see why it is so rare and plus on that Marshosaurus too and I wish they'd find more fossils of Megalosaurus it's been incomplete for so long and it was the first dinosaur ever discovered!
Jeda45 Featured By Owner Jul 1, 2014
What do you think of nasal crests/ridges in megalosauroids? Dennonyx says that it's parsimonious to assume that they had them, since most basal members of "carnosaur-grade" (Ceratosauria, Allosauroidea, Tyrannosauroidea) lineages had them. I'm especially interested in piatnitzkysaurids, so I can make my Marshosaurus reconstruction as accurate as possible.
On a related note, did they have lacrimal horns?
thediremoose Featured By Owner Mar 14, 2014
I'm curious. The specimen of Marshosaurus from the Carnegie Quarry included the back of the skull, the cervicals, and the first five dorsals (which are shown) but also a scapula and partial humerus (which are not). Is the forelimb material no longer considered part of that specimen?
Dennonyx Featured By Owner Mar 23, 2014  Student General Artist
Ezekiel-Black Featured By Owner Mar 11, 2014
Awesome, I love Torvosaurus. Maybe this is a sign for me to do more dino drawings.
ScottHartman Featured By Owner Mar 12, 2014  Professional Digital Artist
It's definitely a sign.
asacquaf Featured By Owner Mar 10, 2014
Where does Edmarka come in on this scale?
ScottHartman Featured By Owner Mar 10, 2014  Professional Digital Artist
Edmarka is just another specimen of Torvosaurus, so its remains have been included already (and the skeletals are not to scale).
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